How plants have adapted or become acclimated to shade. Plants which grow in shady conditions have altered structurally and biochemically in order to cope with the low levels of light and harvest it efficiently. Low level light is rich in far-red light and plants have also adapted to make use of this. Shade plants have also had to make adjustments to cope with sudden high levels of light, for example during sunflecks. Whilst some plants show physiological adaptation, many changes are the result of genetic evolution. Leaves are thinner with a shallow layer of palisade mesophyll cells and a patchy spongy mesophyll with air spaces (fig 1.). This uses less energy and

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